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Well there will be evidence of relevant skills and experiences in the CV, and in the covering letter, you could use the 'and why' part of the third point to mention relevant experience again. The rule though is always do what the employer asks you to do, and if it doesn't make sense immediately try and work out why they are asking for odd or unusual things. Rest assured employers like this will have done quite a bit of thinking about the wording of job adverts. In this case I suspect this employer know it will generally only appeal to people with a relatively strong academic background, so taking that as read, what they are interested in is how people go on and use the opportunity they are offering in future careers. The Wellcome trust is fundamentally a grant giving charity - quite an unusual organisation and it gets high quality applicants for all its jobs. It knows it sets people up for good careers in fund raising, philanthropy, research funding etc and it wants to know people will use that opportunity. So answer their questions, but take any opportunity to give evidence from your life of what you are saying in your answers.